Yeah the usual pronunciation is /ˈdɒ.mɪ.nənt/ or /ˈdɑ.mɪ.nənt/, never /ˈdɑ.mɪˈ
nænt/. But there is still clearly a difference between that and /ˈdɒ.mɪ.nət/, missing the /n/. The difference is probably subtle enough so if you already think it's spelled the other way, you won't hear the distinction.
Etymologically speaking, you would expect the -ant ending for a present active participle and the -ate ending for a perfect passive participle. So something "dominant" would be dominating other things, while something "dominate" would be dominated by other things. Obviously this rule doesn't really work all of the time, but in this case it does.
Grop wrote:But then what is an evolution professor?
A professor of evolution . . . I'm taking a class in Human Evolution and my professor made a point about the spelling of that word because I guess it annoys her.